Threshers are squid-like creatures that like to live in areas with soft ground, as they will burrow in and out of it. They will use their many tentacles to attack players, and come in many different varieties. Their critical hit spot is the tiny eyes on the sides of their head, with the same spots being featured on their tentacles. Note that they cannot be Phaselocked by Maya, instead receiving damage, as they are firmly rooted into the ground.
As one of the weaker kinds of Threshers, they like to either sit back and fling projectiles at you or hit you with their tentacles. They may occasionally burrow underground and leap to another spot.
One of the bigger Threshers, they have more tentacles than the others and will drive them into the ground, popping them back up to attack you. They can also toss projectiles at you if you're too far from them.
Turf Threshers are the largest out of the non-boss Threshers and have their many tentacles being individual targets, instead of counting as one unit. Each tentacle can toss projectiles at you, as well as perform a melee attack if you get too close. You can destroy each tentacle, which deals damage to the main body.
These enemies can be nasty little buggers, as they have a gravitational pull attack that will bring a player to them (similar to a Singularity grenade). Once they do this, spikes will come out of their body and deal constant damage to the player until they either let go or the player deals enough damage.
Badass Pyre Thresher
Similar to a Turf Thresher and with the same attacks, the Badass Pyre Thresher is encased in fire, meaning that it resists incendiary weapons and cannot be set on fire. One new attack it does that other Threshers don't have is that whenever it burrows into the ground and resurfaces, it will release a nova shockwave.